Makino Tsukushi, a working-class girl, attends an elite school called Eitoku Academy (英徳学園, Eitoku Gakuen ?), populated by children and older youths from rich, high-society families. She is the "weed" of the school, surrounded by all the rich kids, including the "Flower Four" (F4). The F4 leader and son of the wealthiest, most powerful family in Japan, Domyouji Tsukasa, takes an interest in Tsukushi, because she is the only girl at Eitoku who does not fawn over him. However, his hot-headed nature and bullying ways are originally a major turn-off for Tsukushi, who has her sights set on someone else (Hanazawa Rui).
The violinist Hanazawa Rui, Tsukasa's best friend, becomes Tsukushi's first serious romantic interest. He is a quiet and cool guy, but he has a soft spot for his close friend and Tsukushi's idol, the model Todou Shizuka, whom he harbored feelings for since childhood. His character is a bit complex, and always has changing feelings for Tsukushi, but above all cares about her a lot.
The other two members of the F4 are Mimasaka Akira, the laid-back peacemaker of the group, and Nishikado Sōjirō, an unrepentant playboy. They both usually have at least one girlfriend at any one time; Akira prefers older women because the women of his household (his mother and two younger sisters) are quite childish. Sojiro is happy to be in casual relationships with many women, although we later discover that at one time he was in love with a childhood friend.
Over the course of the series, however, Tsukushi's feelings evolve, and she begins to appreciate the degree of change that occurred in Tsukasa once he fell in love with her. He becomes fiercely loyal to Tsukushi and believes in her beyond all doubt, and slowly gets her to see him in a different light. The physical obstacles and emotional challenges of their rocky high-school courtship form the basis of the story. Other themes include Tsukushi's attempt to fit in at the school, the problems of her family's lifestyle and income, and the decadent lifestyle of upper-class Japanese girls.
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